Eupatorus gracilicornis

  • Created by: Melissa Phachanhla
  • Completed: April 2015
  • Media:Watercolour, Carbon Dust, Adobe Photoshop, Adobe Illustrator
  • Client:Prof. Dave Mazierski, Dr. Shelley Wall

Project Info:

The images were created from dried out models of the rhinoceros beetle.

Research was compiled on the beetles' living environment.

  1. A.P. Moczek. “On the Origins of Novelty and Diversity in Development and Evolution: A Case Study on Beetle Horns,” Cold Spring Harbor Symposia on Quantitative Biology VLXXIV (2009): 289-296
  2. Armin P. Moczek, Teiya Kijimoto, Emilie Snell-Rood, Guilherme Rocha, Melissa Pespeni, and Karen Kadafar, “Evolutionary and Cological Genomics of Developmental Plasticity: Novel Approaches and First Insights From the Study of Horned Beetles,” Advances in Experimental Medicine and Biology 781 (2014): 127-148.
  3. Devon Troutman, “Rhinoceros Beetle,” last modified Monday, November 14, 2011,
  4. G.J. Arrow, “A contribution to the classification of the coleopterous family Dynastidae,” Transactions of the Entomological Society of London 2 (1908): 321-358.
  5. J. Mark Rowland, Kelly B. Miller, “Phylogeny and systematics of the giant rhinoceros beetles (Scarabaeidae: Dynastini),” Insecta Mundi 0263 (2012): 1-15.